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 See, for example, Juan Comas, "Caracteristicas fisicas de la familia linguistica Maya," Universidad Nacional Autonoma de Mexico, Serie Antropologica 20 (Mexico: UNAM, 1966). Comas compared the results of more than half a century of study of Maya-speaking groups to find marked biological differences among distinct groups within the language community, apparently due to intermarriage with non-Mayan groups, genetic drift, endogamy, and adaptive selection. See also M. Layrisse, Z. Layrisse, and J. Wilbert, "Blood Group Antigen Studies of Four Chibchan [-speaking] Tribes," American Anthropologist 654 (1963): 36-55; the tribes do not form a homogeneous genetic group.